If only one tank is leaking, there will be more water in that tank, water from the bottom of that tank up to the outlet pipe, and half the amount greater than fills up to that outlet pipe, with the other half flowing through the connecting pipe to the other tank.
Unlikely that both tanks would present as leaking at the same time IMHO.
If I've understood this correctly, yes, but I'm not sure what you mean by "fills up to that outlet pipe".
But interconnected tanks will certainly syphon both ways.
F'rinstance, assume there's, say, one pint of water accumulated in the left tank while the car has been parked over night.
(OK, technically, it'll start to syphon immediately the rain water starts to get in.)
If you now park the car on the right side of the road, you'll find the water has syphoned itself to the right tank. This assumes there's a reasonably steep crest in the road.
Similarly, if you then move the car to the left hand side of the road (facing the same way) you'll find the water has moved back to the left hand tank.
I recall going to a BBQ somewhere and parking on a slope. When I later stepped into the car to go home, I noticed the fuel was low but rose back to normal along the way.
There's also "Water Finding Paste". Our guys used this stuff to keep track of water ingress to an underground fuel storage tank we had, back in a past life.
Here's a link to an Oz product (and demo) but it must be fairly common.
https://www.fuelequipmentspecialists.com.au/water-finding-paste/