Well, I'm going to throw in my two cents.
Virtually all engines flowed coolant Up from the block to the head(s) until pretty recently.
Then someone decided that engine temps would be more evenly distributed if the hottest component (head) was cooled first.
I believe Chevy calls this "Reverse flow" and now uses it extensively.
As so many things automotive today this has a lot to do with quicker warm-up for emissions systems.
Also some consideration of complete engine thermal characteristics since so many modern engines use more aluminum and even plastic components than before.
As I recall a double cross-flow radiator is about 40% more efficient for a given size.
So Lotus may have been well ahead of the curve here.